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Transitory and Storage…
Transitory and Storage Starch
Types of Starch: Starch synthesized in leaves directly from photosynthates during the day is categorized as transitory starch, which is later degraded during the night to sustain metabolism and energy production https://diabetesontario.org/flakka-unveiled-navigating-the-chemical-landscape in the absence of photosynthesis. On the other hand, starch in non-photosynthetic tissues, such as seeds, stems, roots, or tubers, is regarded as storage starch and is stored for longer periods to support germination, sprouting, or regrowth when photosynthesis cannot meet the demand for energy and carbon skeletons.
Carbon Partitioning and Regulation
Carbon Fixation and Partitioning: During the day, carbon dioxide is fixed in leaf chloroplasts, and photosynthates are partitioned to satisfy immediate respiratory or sucrose export requirements, with a portion transiently stored as starch in anticipation of the night or during periods of carbon excess.
Regulation of Starch Synthesis: Starch synthesis in the light is tightly controlled in relation to the forthcoming dark period, and starch reserves are still laid down in plants growing under carbon limitation. The production of ADP-glucose by ADP-glucose pyrophosphorylase (AGPase) is the first committed step of starch biosynthesis, and its regulation is crucial for the balance between photosynthesis and sucrose synthesis in leaves.
Role of Enzymes
Enzymatic Steps: Starch synthesis involves several committed enzyme steps, including ADP-Glc pyrophosphorylase, starch synthase, starch-branching enzymes, and starch debranching enzymes, which collectively contribute to the synthesis of starch in photosynthetic and non-photosynthetic organs.
Regulation and Fine-Tuning
AGPase Regulation: AGPase, a key enzyme in starch synthesis, is activated by 3-phosphoglycerate (3-PGA), an intermediate in the Calvin cycle, and inhibited by inorganic phosphate (Pi). The balance between 3-PGA and Pi plays a crucial role in fine-tuning the partitioning between sucrose and starch synthesis, allowing for the regulation of photosynthetic carbon assimilation and starch biosynthesis.
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